Fossil foot bones from A. afarensis add detail to what we’ve learned from the Laetoli footprint trails. For example, Carol Ward, Bill Kimbel and Don JohansonWard:metatarsal:2011 described one of the bones of the midfoot from Hadar, Ethiopia, that represents A. afarensis. This bone, the fourth metatarsal, is the one that connects the fourth toe to the bones of the ankle. In humans, these ankle bones are higher, and transfer the body’s weight downward into the arching midfoot. So the fourth metatarsal has to be slightly twisted as it arches down toward the lateral (outside) side of the foot. A chimpanzee’s foot is much flatter, so the bone doesn’t twist. Ward and colleagues found that the A. afarensis bone was twisted in a humanlike way.