Consider - an evolutionary advantage means reproductive success; when a population evolves towards niche construction, what is the process that selects for the relevant behaviors?
As I have already shown in the previous post that it is an assumption to say that natural selection alone is the process that selects for relevant behaviors. Even before natural selection comes along it is the creature that is selecting the healthy and positive conditions that will enhance their fitness and provide benefits for their offspring in future generations. So long as that niche is maintained it will provide heritable benefits.
When a population of niche constructors reproduces, generating variant offspring, what is the process that selects for those variants that maintain or improve on those behaviours? IOW what is the process which establishes whether a certain pattern of behaviour is advantageous?
The process that establishes whether a certain pattern of behavior or environment is advantageous is the creature itself through changing its environment, its developmental responses to its environment, and the back and forth feedback between these factors as well as NS. Natural selection is not the privileged or only force at play here and that is what the EES is trying to dispel. I think I have provided ample evidence for this in the other responses. The fact that you still want to emphasize selection over everything else shows even though you claim that the SET acknowledges the EES forces they don’t really recognize them as actual causes that direct evolution like NS.
Niche construction can be summed up here
However, niche construction is not restricted to the biological adaptations of organisms but also stems from their developmental plasticity, their by-products, and their acquired characters. Organisms are viewed as active agents that impose order on the conditions of their existence, and direction on their development and evolution.
Niche construction theory emphasizes how niche construction can scale-up across individuals in populations and overtime to generate a stable and directional modification of environmental components. As a result, niche construction generates consistent and sustained forms of natural selection and thereby impose biases on evolution (Laland et al, 2015).
Niche construction
As the article points out “niche construction generates consistent and sustained forms of natural selection”. Creatures have the ability to create conditions that provide well-integrated and suited changes that provide fitness and adaptive benefits just like NS can do and therefore is another source that determines the course of evolution and in doing so biases and diminishes NS.
Which behaviours in which creatures, for example?
As far as niche constructions many creatures such as insects, birds, and mammals, basically most creatures that produce offspring create conditions conducive for healthy birthing and rearing through a number of behaviors. These include creating protective birthing conditions with insects such as the Dung Beetle or with birds and mammals.
As far as inheritance beyond genes we now know through processes like epigenetics and direct interactions that whatever behaviors a creature has and teaches their offspring will affect and influence future generations. We know that social and cultural practices can help with evolvability and put creatures in better positions to thrive and survive. These forces are part of the EES and influence the course of evolution by helping creatures be in a more adaptive position.
What, exactly, do you mean by learnt behaviour? Learnt how - by imitation? It can't be trial and error.
Under inheritance beyond genes creatures' behaviors can be passed along and down to future generations through social and cultural practices which can be advantageous similar to niche construction in putting creatures in a better position to be fitter and adapt. These behaviors like niches can be passed on and provided the creatures maintains that behavior or niche and adjusts it accordingly when needed it will continue to provide benefits now and in the future. But behavior can also cause negative outcomes as we have seen with humans.
I think most animals have an inherent ability to work with nature and therefore their behaviour is more likely to provide benefits. But humans often work against nature as well. But we also have the ability to make a big difference in rectifying the harm done and counteracting those negative outcomes by reestablishing nature and creating artificial environments that help cultivate environments and niches. As we have seen humans are so good at mitigating natural consequences such as through medicine and technology that we are overriding NS influence altogether. To a lesser extent, all life has this ability but to a lesser extent as mentioned with niche construction and inheritance beyond genes.
You say niche construction behaviours "always produce advantageous results", but it's common that the predictability of niche behaviours allows predators and parasites to specialise on them, and in the case of literal niches, for usurpers to requisition or 'cuckoo' the niche. When environmental conditions change, a niche may become uninhabitable, unconstructible, or may simply disappear, leaving the niche constructer specialised for a non-existent or unconstructible niche... In what sense are these 'advantageous results'?
I am not sure it is common and the fact that this happens doesn’t negate the important role niche construction plays as a cause of evolutionary change. According to NCT creatures are not dumb passive creatures that cannot change with changing environments. They have a great capacity to adjust environments to suit. Living things seem to have a ready set of behaviors and tools available (hormonally, chemically, etc.) that help create the right environment. It seems inherent in them.
They are in tune with nature and their particular environment, what the dangers are, and how to set things up in the best possible way to grow, develop, and survive. That’s not to say that mistakes are not made and disasters don’t come along or that creatures are just as smart in getting around these niches. But these may be individual occurrences for both creatures and environments. Each will be different and are not the environment or species-wide events. If creatures are found out then they have the capacity to learn and try to create a better niche.
But this is different from the narrow view, SET takes that treats creatures as passive entities subject to predators and changing environments where they can only survive if they are adapted genetically through NS.
Developmental plasticity only gives a limited range of adaptability; it will not always produce 'precisely what is needed' for an environment. Random mutations still occur, but they are relevant over multiple generations, whereas developmental plasticity is an individual response.
As far as I understand there is a certain scope for plasticity through development just like there is a certain scope of variation developmentally in the first place. There is good reason for this as only certain variations work in environments so saying that random mutations are going to find a better solution that doesn’t seem to be supported.
Plasticity is often the first step in adapting to environments and usually produces well suited and integrated change that may be cemented later with NS. But it doesn’t take generations and can happen pretty quickly because there is only certain variations that can be produced and because taking the constructive and reciprocal view of the EES creatures are not separate to their environments and evolve with their environments. So variations are often developmental responses to the environments they are in.
What is the process that selects for developmental plasticity over developmental fixity/rigidity? What is the process that selects for a particular type of developmental plasticity in a changing environment?
When you consider that there are only certain variations that can be produced and only a certain scope within those variations, it seems to me that initially and primarily it is the development system that is responding to environments and producing the required changes as a result. As mentioned above this seems to be the main way that development works and NS is something that may come along later and refine and fix these plastic changes in.
But when you consider that these developmental changes are unlike the random ones from mutations but are nonrandom then they are more directed at achieving specific changes that have been determined to be most suitable. This would mean that it is not NS but developmental systems that are determining which variations work best and not NS or at least not NS on its own but both working together.
Good grief... that is '
not even wrong'. It reflects a fundamental lack of understanding or joined-up thinking about evolution.
Why I can support everything that is said here. I think I have already covered this above. For example, how the SET treats the environment and creatures as separate entities whereas the EES sees them intertwined and therefore affecting each other is supported here
Central to these debates are different assumptions concerning the independence or interdependence of the causes of phenotypic variation, differential fitness, and inheritance, which are Lewontin’s (1970) three conditions for evolution by natural selection (Walsh 2015; Uller & Helanterä 2019). Traditionally, evolutionary biologists have assumed these processes are quasi-independent, but in practice, they are often causally intertwined.
Niche construction
As far as showing that random mutations that produce random variations not being conducive for advantageous changes and therefore need to be tested and sifted by natural selection that's easy to explain without any references. As random mutations cannot be predetermined as to whether they provide a benefit for adaptations they need NS to determine their worth by testing each variation against the environment.