For those that would like more of a basic overview and how it fits in with the rest of evolutionary thinking, check out the Wikipedia article.
Extended evolutionary synthesis - Wikipedia
Thankyou Kylie. This is a pretty good explanation of the EES. Just to point out a couple of things mentioned. Under predictions
Prediction 1. Change in
phenotype can precede a change in
genotype[4]. So evolution can be non-gene based as with the traditional theory.
Prediction 2. Changes in phenotype are predominantly positive, rather than neutral (or negative my emphasis).
As phenotype change can be the result of self-organization where living things have a say in what happens to them and the result of feedback from other living things and the environment acting through development processes changes to phenotypes are often well suited to environments and integrated into existing genetic networks. This is opposed to the adaptive view under the traditional theory where mutations are random and creatures have to be passively adapted to environments through selection.
Prediction 6. adaptation can be caused by natural selection, environmental induction, non-genetic inheritance, learning, and cultural transmission.
This is supported by what is mentioned in prediction 2. Natural selection is only one of a number of evolutionary influences and not any more dominant than the others especially when it involves complex genetic networks whose structure and integrity need to remain the same and avoid random mutations that can undermine those structures.
Prediction 9. Heritable variation is directed towards variants that are adaptive and integrated with phenotype
[4
It seems to me that rather than any variation being thrown up for natural selection as with random mutations variations seem to be very suitable not only to the environment but integrated into the creatures system in the first place without going through a filtering process of finding the beneficial changes among many other alternatives.
Unlike the so-called beneficial mutations which are really not a complete benefit as they often have a trade-off in knocking out some function, the adaptive and integrated changes through these other processes seem to be a perfect fit. This makes it much easier for selection and in fact is more or less doing selections work.
Prediction 10.
niche construction is biased towards environmental changes that suit the constructor's phenotype, or that of its descendants, and enhance their fitness
[2]
As with the above, these changes can be very well suited and bypass the need for selecting out dysfunction and weak as living things are able to have some control over their environments. Another process overlooked where natural selection can be given credit whereas other forces are at play.