None of that says, "natural selection will be diminished, biased, and even replaced by the EES forces".
Then you do not really appreciate what the paper is implying. If under the SET Natural selection is seen as the only mechanism for causing and directing evolution then it logically follows that including several other modes of transmission that do the same thing would then take away from natural selections role in some way whether that be biasing, diminishing, and replacing NS in different situations.
Biasing the outcome of natural selection is not biasing natural selection; any consistent change in selection pressure will 'bias' the outcome of natural selection.
But this doesn't make much difference when we are talking about natural selection directing the outcome of evolution. Also, there is evidence that natural selection's role itself is being affected by other forces that can also act as selecting forces. Therefore they are more or less replacing or reducing NS as a result IE
There is another process of potentially substantial evolutionary influence: the (constructed) environment does not just select for new variation, it also produces it, in the form of the developmental niche.
Why developmental niche construction is not selective niche construction: and why it matters
So as you can see EES forces like niche construction can not only produce nonrandom variations but also select those adaptive variations therefore taking over the role that NS would usually do and in doing so reducing it.
There's no mention of natural selection being diminished or replaced.
As mentioned above you are not appreciating the implications of what the papers are saying about how the EE forces can do what NS does. If the SET claims that NS is the sole force that causes and directs adaptive evolution by being the only selecting force and the EES forces can do what NS can do then it logically follows that NS role will be reduced and diminished as a result. There is evidence that for example, niche construction also acts as a selecting force thus replacing NS or at least diminishing its role by becoming the force that determines adaptive variations instead of NS.
We suggest that niche construction occupies the middle ground between artificial and natural selection: like artificial selection, niche construction reliably generates relatively consistent features in selective environments. During artificial selection, breeders and experimentalists deliberately select for particular characteristics (high yields, pretty flowers and attractive plumage); the breeder/experimentalist imposes direction on evolution by determining which individuals reproduce. There is a predictability and consistency to the pattern of evolution that ensues—the breeder/experimentalist can anticipate with confidence that a specific favored trait will reliably increase in frequency until genetic variation is significantly depleted and can predict with some accuracy the direction of evolution. Selective breeding increases the frequency of the selected trait, frequently evoking characteristic and strong responses to selection.
The predictability and generality of artificial selection can be contrasted with the frequent unpredictability and local contingency of natural selection in natural populations without niche construction.
https://royalsocietypublishing.org/doi/10.1098/rsfs.2016.0147
Our study confirmed that natural selection deriving from organism-constructed sources exhibits reduced temporal and spatial variation in selection gradients, and weaker (i.e. reduced intensity of) selection, compared to non-constructed sources.
Organism-Constructed Environments are Different – Extended Evolutionary Synthesis
Nope. You're just repeating the same old nonsense. Also, natural selection doesn't 'come along', it is always acting.
I think I have supported each claim above. I won't go through all of them but for example
I said
“Under the SET view the source of variation is random so natural selection is regarded as the all-important and only determining factor as to which variation are adaptive and heritable. Therefore NS is seen as the driving force of evolution.”
Let’s see what the papers say.
Like the EES, the Modern Synthesis also represents a particular way to understand evolution. It primarily focuses on genes:
- new variation arises through random genetic mutation
- inheritance occurs through DNA
- natural selection of genes is the sole cause of adaptation
http://rspb.royalsocietypublishing.org/content/282/1813/20151019
EES processes can't 'replace' natural selection - no natural selection means no evolution...
If the EES forces are not replacing NS then how do you explain the above quotes from papers saying they do.
Natural selection is not a theory, it's a process;
Then why do so many articles refer to it as a theory IE
The theory of natural selection was explored by 19th-century naturalist Charles Darwin. Natural selection explains how the genetic traits of a species may change over time. This may lead to speciation, the formation of a distinct new species.
Natural Selection.
conceptually different from niche construction. If there is selection, it is natural selection; it may be considered as part of a greater process, but it acts just the same.
Then how do you explain niche construction for example being said to also be a selective force that produces and selects adaptive variations.
None of that supports what you said above - in fact they explicitly say, "Causation in biological systems is reciprocal rather than linear, with natural selection and niche construction codirecting each other."
That exactly supports what I said. I said the SET takes a narrow view of the cause of evolution which is linear. Whereas the EES takes a more pluralistic view where there are a number of forces that cause and drive evolution. I have never denied NS is one of them, just that it is not the only one like the SET claims and just that the EES forces also cause and drive evolution which SET denies.
This is simply multi-generational kin selection - where altruistic actions by particular population variants with a mutant allele benefit not only their existing normative kin, but, by extension, those of future generations. By benefitting kin survival, the mutant allele (particularly recessive alleles) can persist over the generations, benefitting future generations.
Actually it is more then this, more than altruistic behaviour and more than just passing on alleles. Any behaviour or niche conditions that may provide an advantage are preserved regardless of NS, regardless of whether that specific condition of behaviour is a selective or adaptive benefit according to Natural selection ad regardless of whether it is gene-based or not which the SET restricts things too.
As shown it is the creature’s actions and the niches that are doing the selecting and this will determine adaptiveness and heritable variations according to what the creature itself requires which may not always be the same as the adaptive and fitness to environments that natural selection determines. So any conditions and behaviours that provide this adaptive benefit despite NS will be passed on if it helps the creature and future generations survive. IE
Niche construction processes can facilitate the endurance of organisms in adverse environments and it can be beneficial despite being costly due to advantages that accrue for later generations [99].
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Note that the cause here is a mutation and it is subject to natural selection.
As mentioned the EES doesn't restrict variations that provide an adaptive and survival benefit to gene-based variations such as with inheritance beyond genes.
Niche construction is the organism adapting to its environment by changing its behaviour to modify the environment. As previously explained, the difference is just a matter of viewpoint and categorization.
Niche construction is not as you describe it where an organism is adapting to its environment. As soon as you say this you are still taking the SET adaptive view which restricts all-cause to adaptive evolution by NS. Rather it is the environment being adapted to the creature (the other way around) then the creature being adapted to the environment.
This makes an important distinction. Saying the creature is adapted to its environment still makes NS the cause of that adaptation. The outside force of NS bearing down on some passive creature who is being changed to fit that environment. It overlooks any contribution from the creature itself or its developmental system that may be doing the selecting and therefore controlling its own evolution.
Under Niche construction, the creature has changed the environment to suit its needs. As I have shown above niche construction is a form of selection itself. So instead to NS doing the selecting of adaptive variations, it is the creature through the niche conditions, it has created that will suit it now and in future generations to help it adapt and survive meeting the conditions for evolution.
This is standard ecosystem interaction; nothing new here. EES simply describes it from a different perspective.
Yes and as I have just shown above that different perspective is important. It brings with it different scientific assumptions, structures and predictions for causes of evolution. Because the EES takes a more pluralistic view of the evolutionary causes. It takes into consideration other forces that cause and direct evolution and their greater influences and feedbacks besides the adaptive view through natural selection.
The difference is that the SET sees those other forces and greater influences as explanations for why adaptive evolution by natural selection is being constrained or is absent because it only sees all causes as by NS. Whereas the EES sees those EES forces causing and directing evolution and thus the greater influences are part of this expanded view as contributing to the cause of evolution.