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Mas sabe el diablo por viejo que por diablo.
I highly doubt the social animals have any abstract notion of "morality". They may have their specific morality and follow it (usually), but the abstraction is far from their little minds.
I didn't say anything about abstraction or much in the way of self awareness, however this doesn't change the fact that morality is, like anything else in the universe, a continuum that joins the most basic automated chemical responses of ameoba to our complex and often conflicting problems of the individual versus the group.
God is merely an explanation humans came up with to answer their original questions, while these questions may have appeared very early in human evolution, the fact remains that animals, including our own ancestors had been faced with moral decisions and acted upon them probably for hundreds of thousands, if not millions, of years prior.
Just like animals may not have the abstract notion of 'love,' the feelings of affection, care, concern, attraction, etc are all observed in the animals kingdom.
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