Who decides, no one. The outcome is based on whether the genetics or environment was the stronger.
If you ate a chocolate bar every day of your life and then decide not eat it one day, it would be because there is a reason. You can choose to act against your gentics, but that would mean your environment caused you to overcome your genetics. You can choosed to act against your environment, but that would mean your genetics overcame your environment.
I think we're at an impass caused by differences in philosophy. You assume that everything in my consciousness, in other words everything that constitutes myself, is determined by genetics and environment. I on the other hand do not believe this to be the case. For starters, I've already mentioned my ability to make different choices in the same circumstances. My genes are the same from the moment my mom's egg met by dad's sperm up to my death. My environment doesn't vary all that much. In the life of an ordinary math teacher, one school day is more or less like the next. Yet my choices on a given day are often vastly different. Moreoever, I can experience the fact that different choices don't arise from different, prior mental states. I always wake up in the morning feeling the same way: groggy, tired, and a little bit achy. Yet even so, I can make vastly different decisions right off the bat.
How do you choose to act against both simultaneously?
To answer that rigorously, we'd first have to know exactly what genetic and environmental factors influence my thinking. And that would be difficult. Because, believe it or not, scientists have not yet found
any gene that has
any effect on human thinking. There have been a lot of large, thorough scientific studies
trying to locate such genes, but in all cases there are no results consistent enough to be sure of the fact. Similarly there are attempts to explain human thinking based on chemical influences such as serotonin, adrenaline, epronepron, and so forth. However in recent years those theories have been put on shaky ground by scientific studies.
However, I can say this much, that in some cases where I feel the (alleged) adrenaline rush and desire to run around or jump or doing something else physical, I'm still able to stand still. In cases when my (alleged) low serotonin levels make me want to lie down and mope for a while, I'm still able to get up and run and jump. And so forth.
While, it is true that you can overcome your genetics as a result of your environment; and it is true you can act differently then your past decisions that may have been the result of your past environment. You still acted as a result of your current(up to the moment) environment(i.e. thoughts leading up to the moment). There was a sequence of thoughts that took place that lead to the decision. All cause and effect.
As for the experiment in the video, it's worth noting that Dr. Benjamin Libert of UCSF, the first neurologist to perform that experiment, specifically said that the results did not rule out free will.