The article is open access, thus there is no reason why you you should not be free to discuss it giving proper citation. In fact, you can download it for your own private use. What you did was copy and paste the entire article making it available outside of NCBI or designated sites to display it, which is forbidden and stated so on the NCBI site.
As for your comment about his quote on the origin of life research, I couldn't find any such statement. He does mention the origin of species several time, which is the title of Darwin's work. And I saw nothing about Koonin putting down evolution. All he did was provide a bit of history of what has been learned at the centennial of the publication and what has been learned between the 100 yr. and 150 yr. mark. He even states that the next 50 yrs. will show even more developments and changes of understanding.
I would suggest messaging the moderator who discussed this with you and ask for clarification, and cite my posts concerning this discussion if you like. I certainly don't know, but I think there may be a misunderstanding on what the moderator was conveying, but for sure, posting the entire article here was a really big no no. Good luck.