I've seen it raised a few times on this forum that natural selection is metaphysics. I challenge that claim with the following:
First of all, I snagged a definition of metaphysics from Dictionary.com:
met·a·phys·ics ( P ) Pronunciation Key (mt-fzks)
n.
1. Philosophy. The branch of philosophy that examines the nature of reality, including the relationship between mind and matter, substance and attribute, fact and value.
2. The theoretical or first principles of a particular discipline: the metaphysics of law.
3. A priori speculation upon questions that are unanswerable to scientific observation, analysis, or experiment.
4. Excessively subtle or recondite reasoning. (modified for readibility)
Of these, definitions two and three are where natural selection might fall under relating to biological evolution. But definition 2 is vague, whereas definition 3 is precise: A priori speculation upon questions that are unanswerable to scientific observation, analysis, or experiment.
Second, I snagged a definition of natural selection, also from Dictionary.com
n.
The process in nature by which, according to Darwin's theory of evolution, only the organisms best adapted to their environment tend to survive and transmit their genetic characteristics in increasing numbers to succeeding generations while those less adapted tend to be eliminated.
Now, if this process is "a priori speculation upon questions that are unanswerable to scientific observation, analysis, or experiment", therefore it should not be possible to test (scientifically) natural selection.
However, I have an example in which the outcome of the process of natural selection was predicted and verified via a laboratory experiment:
Multiple Duplications of Yeast Hexose Transport Genes in Response to Selection in a Glucose-Limited Environment
Excerpts:
The Evolved Strain Outcompetes the Parental Strain when they are Grown Together in Continuous Culture
Previous observations showed that the evolved strain had reverted to the GAL1 phenotype; 28–15L4 and CP1AB are therefore readily distinguished by colony size on 0.8% galactose minimal agar. A pair of chemostats was initiated with equal densities of the parental and evolved strains, and their relative frequencies were followed for 20 generations (fig. 2). The frequency of the evolved strain increased steadily in both chemostats until the parental strain could no longer be detected.
The Evolved Strain Transports Glucose Two to Eight Times Faster than the Parental Strain
Given our observation that the two strains differ in s by an order of magnitude, the simplest explanation for their difference in competitive ability is that selection has favored the evolution of an improved mechanism for transporting limiting substrate. Figure 3 shows the results of glucose transport assays comparing uptake velocity at several glucose concentrations for cells grown in chemostat monoculture on 0.08% glucose at a dilution rate of 0.2/h. The evolved strain consistently demonstrates greater substrate uptake velocity than the parental strain.
The full paper goes into much more detail, but I highlighted the parts I need to make my point. We have an empirical demonstration of natural selection in action, with a particular strain of yeast out-competing another in response to its environment.
Is natural selection metaphysics? Based on the above experiment, I would say no. So to the creationists that have made this assertion, can you defend it? If this experiment doesn't demonstrate natural selection, then what are we seeing here?
First of all, I snagged a definition of metaphysics from Dictionary.com:
met·a·phys·ics ( P ) Pronunciation Key (mt-fzks)
n.
1. Philosophy. The branch of philosophy that examines the nature of reality, including the relationship between mind and matter, substance and attribute, fact and value.
2. The theoretical or first principles of a particular discipline: the metaphysics of law.
3. A priori speculation upon questions that are unanswerable to scientific observation, analysis, or experiment.
4. Excessively subtle or recondite reasoning. (modified for readibility)
Of these, definitions two and three are where natural selection might fall under relating to biological evolution. But definition 2 is vague, whereas definition 3 is precise: A priori speculation upon questions that are unanswerable to scientific observation, analysis, or experiment.
Second, I snagged a definition of natural selection, also from Dictionary.com
n.
The process in nature by which, according to Darwin's theory of evolution, only the organisms best adapted to their environment tend to survive and transmit their genetic characteristics in increasing numbers to succeeding generations while those less adapted tend to be eliminated.
Now, if this process is "a priori speculation upon questions that are unanswerable to scientific observation, analysis, or experiment", therefore it should not be possible to test (scientifically) natural selection.
However, I have an example in which the outcome of the process of natural selection was predicted and verified via a laboratory experiment:
Multiple Duplications of Yeast Hexose Transport Genes in Response to Selection in a Glucose-Limited Environment
Excerpts:
The Evolved Strain Outcompetes the Parental Strain when they are Grown Together in Continuous Culture
Previous observations showed that the evolved strain had reverted to the GAL1 phenotype; 28–15L4 and CP1AB are therefore readily distinguished by colony size on 0.8% galactose minimal agar. A pair of chemostats was initiated with equal densities of the parental and evolved strains, and their relative frequencies were followed for 20 generations (fig. 2). The frequency of the evolved strain increased steadily in both chemostats until the parental strain could no longer be detected.
The Evolved Strain Transports Glucose Two to Eight Times Faster than the Parental Strain
Given our observation that the two strains differ in s by an order of magnitude, the simplest explanation for their difference in competitive ability is that selection has favored the evolution of an improved mechanism for transporting limiting substrate. Figure 3 shows the results of glucose transport assays comparing uptake velocity at several glucose concentrations for cells grown in chemostat monoculture on 0.08% glucose at a dilution rate of 0.2/h. The evolved strain consistently demonstrates greater substrate uptake velocity than the parental strain.
The full paper goes into much more detail, but I highlighted the parts I need to make my point. We have an empirical demonstration of natural selection in action, with a particular strain of yeast out-competing another in response to its environment.
Is natural selection metaphysics? Based on the above experiment, I would say no. So to the creationists that have made this assertion, can you defend it? If this experiment doesn't demonstrate natural selection, then what are we seeing here?