What Darwin was defending against was this:
"But to the progenitor of the upland goose and of the frigate-bird, webbed feet no doubt were as useful as they now are to the most aquatic of existing birds. So we may believe that the progenitor of the seal had not a flipper, but a foot with five toes fitted for walking or grasping; and we may further venture to believe that the several bones in the limbs of the monkey, horse, and bat, which have been inherited from a common progenitor, were formerly of more special use to that progenitor, or its progenitors, than they now are to these animals having such widely diversified habits.
Therefore we may infer that these several bones might have been acquired through natural selection, subjected formerly, as now, to the several laws of inheritance, reversion, correlation of growth, &c. Hence every detail of structure in every living creature (making some little allowance for the direct action of physical conditions) may be viewed, either as having been of special use to some ancestral form, or as being now of special use to the descendants of this form either directly, or indirectly through the complex laws of growth.
Natural selection cannot possibly produce any modification in any one species exclusively for the good of another species; though throughout nature one species incessantly takes advantage of, and profits by, the structure of another. But natural selection can and does often produce structures for the direct injury of other species, as we see in the fang of the adder, and in the ovipositor of the ichneumon, by which its eggs are deposited in the living bodies of other insects. "
Darwin was saying that structures produced in one creature were not "exclusively" for the benefit of one of it's ancestors.
He would be shocked to see that structures would be produced for the "exclusive" benefit of a totally unrelated organism.
It would be like Hume, after telling Paley that watches and organisms were too dissimilar for a good analogy, seeing molecular motors. These have one to one correlations to man made motors.
So, I think the links that I provided earlier, falsify Darwins theory.
"If it could be proved that any part of the structure of any one species had been formed for the exclusive good of another species..." and in these cases totally different species.
God Bless!
And good night.